In the context of current concerns within the environmental humanities to challenge the idea that humans are somehow irreducible to nature, this article takes up the much-neglected history of the idea of human exceptionality itself. According to now familiar accounts, metaphysical assumptions about the unique status of the human are considered to have persisted-including to the present day-despite evolutionary contentions that the human should be understood as a purely physical being. Such, largely Christian and Cartesian, metaphysical notions of a human soul or mind doubtlessly endure. But in this article we consider the-largely ignored, yet now arguably more prevalent-idea that humans are exceptional because of their physicality. Here, th...